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Chiltern Railways is a British train operating company that provides passenger services between London Marylebone and Birmingham Snow Hill, as well as other destinations in the Midlands and South East England. The company was established in 1996, and it is owned by Arriva UK Trains since 2009. Chiltern Railways operates a fleet of modern trains, including the Class 68 locomotives and the Class 165 and 168 diesel units.

East Midlands Railway is a train operating company that provides passenger services across the East Midlands region of England. It was formed in 2019, taking over the franchise previously operated by East Midlands Trains. The company operates services on routes including London St Pancras to Sheffield, Nottingham to Liverpool, and Norwich to Liverpool. East Midlands Railway also operates local services within the East Midlands, connecting towns and cities such as Derby, Leicester, and Lincoln.

London North Eastern Railway (LNER) is a train operating company in the United Kingdom that operates intercity services on the "East Coast Main Line" between London, Yorkshire, the North East and Scotland. It was formed in 2018 as a result of the UK government's decision to terminate the franchise of Virgin Trains East Coast. LNER is owned by the Department for Transport and managed by the operator of the East Coast Partnership, a joint venture between infrastructure company Network Rail and train operator FirstGroup. LNER operates a fleet of electric and diesel trains, including the Azuma, which is the fastest train in the UK.

London Overground is a suburban rail network in London, England. It was created in 2007 by the merger of the former Silverlink Metro service with other rail services. The network consists of 112 stations and serves 23 London boroughs. London Overground operates on these routes, which include Highbury & Islington to West Croydon/Clapham Junction/Crystal Palace, Richmond/Clapham Junction to Stratford, Gospel Oak to Barking Riverside, Watford Junction to Euston, Liverpool Street to Enfield Town/Cheshunt, Liverpool Street to Chingford, and Romford to Upminster. Overground is operated by Arriva Rail London under a concession from Transport for London.

The London Underground, also known as "the Tube", is a rapid transit system that serves London and some parts of the surrounding counties. It is the oldest underground railway network in the world, with the first line opening in 1863. The Tube is a vital part of London's transportation infrastructure, carrying over five million passengers per day. It is operated by Transport for London (TfL) and is known for its iconic map, distinctive roundel logo, and deep-level tunnels.

TransPennine Express is a train operating company in the United Kingdom that provides intercity train services across the North of England and Scotland. The company operates services between major cities such as Manchester, Liverpool, Leeds, Newcastle, Edinburgh, and Glasgow. Transpennine Express offers a range of ticket options, including advance purchase tickets, out-of-season tickets, and flexible tickets. The company also provides onboard facilities such as free Wi-Fi, power sockets, and refreshments.

West Midlands Railway is a train operating company that provides passenger services across the West Midlands region of England.The West Midlands Trains consortium, composed of Abellio, JR East, and Mitsui, won the franchise for the region, leading to the establishment of the service in 2017. The West Midlands Railway offers a variety of services, including longer distance services to cities like London and Liverpool as well as local and regional trains.